When a domain controller has been offline for more than the specified tombstone lifetime, it is considered bad and will no longer replicate properly with the other controllers.
When this happens, new users, groups, and other objects will not be synchronized anymore on this server. It can cause issues with emails sent to these new users. If the email server can still check for the user in the AD against the bad server, emails won’t be delivered since it thinks that the user does not exist.